Sauropods

Giants of the vineyards

Fossils from Angeac-Charente attest to at least two lineages of herbivorous sauropods: a turiasaur whose remains have been collected since 2010, and a macronarian close to Camarasaurus, discovered in 2024.

Angeac sauropod: reconstruction drawn in April 2013, before the macronarian discovery in 2024.
Angeac sauropod: reconstruction drawn in April 2013, before the macronarian discovery in 2024. © Mazan.
2.20 mthe length of a large femur
Over 30 mthe estimated length of the giant turiasaur
Herbivoresfeeding on plants

Two sauropod lineages at Angeac

The turiasaur, known since 2010

Since the first excavation campaigns, large femora, teeth and many other bones have attested to a turiasaur at Angeac. These fossils are assigned to Turiasauria; their species remains undetermined.

Research on turiasaurs brings together Rafael Royo-Torres and the Teruel-Dinópolis teams. Comparisons between fossils from Angeac and the Iberian Peninsula refine our understanding of the anatomy and evolution of this sauropod group.

Meet Rafael Royo-Torres → · Dinópolis and Teruel →

Turiasaurus: reconstructed skull, head and silhouette. A comparison for understanding turiasaurs, without assigning the Angeac fossils to this genus.
Turiasaurus: reconstructed skull, head and silhouette. A comparison for understanding turiasaurs, without assigning the Angeac fossils to this genus. © Mazan.

2024: the discovery of a different sauropod

Opening a new excavation area exposed the end of a femur. Within three weeks, the excavators had uncovered part of a sauropod skeleton, with several bones still articulated. Initially thought to be another turiasaur specimen, the animal proved anatomically distinct, particularly in its large, spatula-shaped teeth.

The shape of the teeth, the very broad vertebrae with bifid neural spines and the robust pelvic bones led Ronan Allain to identify a new species close to Camarasaurus, within Macronaria. It is therefore a different sauropod from the turiasaur found since 2010.

By the end of the 2024 season, the excavated remains included four articulated dorsal vertebrae, about twenty ribs, three pelvic bones, a femur, much of the lower jaw and around twenty teeth. Initial estimates place its length at 15–20 metres and its mass at about 30 tonnes. Its relationship to camarasaurs suggests a relatively short neck that could be raised to reach higher vegetation.

Proposed reconstruction of the Angeac macronarian sauropod, close to Camarasaurus. Its complete appearance is inferred from fossils and comparisons.
Proposed reconstruction of the Angeac macronarian sauropod, close to Camarasaurus. Its complete appearance is inferred from fossils and comparisons. © Mazan.

Interview with Ronan Allain, MNHN, 20 September 2024 ↗

What the recovered bones allow us to reconstruct

The skeletal drawing places the recovered bones: vertebrae of the neck, back, sacrum and tail, ribs, shoulder girdle, pelvic bones, femur and lower jaw. In Mazan’s plate below, the elements highlighted in red correspond to the illustrated record of the 2024 and 2025 finds. Parts completed through comparison help reconstruct the whole animal.

Reconstructed skeleton of the Angeac macronarian; bones from 2024–2025 highlighted in red. Annotated plate by Mazan, 2026.
Reconstructed skeleton of the Angeac macronarian; bones from 2024–2025 highlighted in red. Annotated plate by Mazan, 2026. © Mazan.

A complete skeletal drawing is not a complete skeleton recovered in the field. The label “Camarasauridae indet.” on the drawing indicates a proposed family-level identification, without a determined genus or species. Similarity to Camarasaurus guides anatomical comparisons; it does not replace the scientific description of the new material.

Lower jaw of the new sauropod, still surrounded by sediment in its protective jacket.
Lower jaw of the new sauropod, still surrounded by sediment in its protective jacket. © Mazan.
Reconstructed skull and lower jaw for the sauropod close to Camarasaurus. Drawing by Mazan, 2024; missing parts are interpreted.
Reconstructed skull and lower jaw for the sauropod close to Camarasaurus. Drawing by Mazan, 2024; missing parts are interpreted. © Mazan.

From a protected block to anatomical observation

The sketch made at the Musée d’Angoulême in March 2025 shows the ischium, a pelvic bone, and ribs in their block. Before comparing these bones, they must be exposed and stabilised while retaining their numbers and field information. The drawing also documents this stage of the work, between excavation and study.

Ischium and ribs of the new sauropod in their block, Musée d’Angoulême, 19 March 2025. Mazan’s sketchbook.
Ischium and ribs of the new sauropod in their block, Musée d’Angoulême, 19 March 2025. Mazan’s sketchbook. © Mazan.

Compare silhouettes and skeletons →

Reconstructing the giant, bone by bone

Teeth, skull fragments, neck, back and tail vertebrae, pelvic bones, limbs and foot bones allow us to compare different parts of the skeleton. The giant sauropod known since the first field seasons belongs to Turiasauria.

Each new element refines our understanding of its anatomy. A claw from the first toe, discovered after a violent storm in 2013, adds another detail to the animal’s reconstruction.

Digital reconstruction of the Angeac environment with sauropods
Digital reconstruction of the ancient environment. © Michel Fontaine, 2015.

An animal in its landscape

We study sauropods as part of their whole environment: rivers, wetlands, vegetation and other animals. Reconstructions show the connections between these discoveries and the habitats in which the giants lived.

Reading the tail, vertebra by vertebra

Caudal vertebrae form the tail. Their sequence reveals repeated structures and changes in shape along this part of the skeleton. Researchers also examine articulation surfaces, arches and bony projections: characters used in comparisons between sauropods.

Anatomy of Angeac sauropods: Allain et al., 2022 ↗

Caudal vertebrae of the Angeac turiasaur arranged for examination.
Caudal vertebrae of the Angeac turiasaur arranged for examination. © Ronan Allain.

A tooth, patiently reassembled

A tooth’s outline may only emerge after lengthy preparation. This specimen arrived from the excavation in many fragments. Sorted and matched one by one at the museum, they gradually fit together until the crown and the beginning of the root become recognisable.

Reassembly reveals the fossil’s shape without erasing its fractures. Outlines, surfaces and missing areas must remain distinguishable for observation and scientific comparison.

Sauropod tooth after reassembly of the fragments: fourth stage of the photographic sequence.
Sauropod tooth after reassembly of the fragments: fourth stage of the photographic sequence. © Mazan.

See this tooth’s reconstruction, step by step →

Research published in 2026

Different teeth in the same jaw

The study of 167 isolated teeth from the Angeac turiasaur identifies five morphotypes linked to their position in the jaw. Comparative anatomy and 3D morphometrics work together to classify teeth and resolve ambiguous cases.

Clémence Le Gouellec, Ronan Allain, Jean Goedert, Romain Pintore, Jackson Carter, Rafael Royo-Torres, Dominique Augier and Raphaël Cornette, Journal of Anatomy, 2026.

Read the open-access study and figures ↗

Specimens and data

From teeth to the giant femur

The diversity of teeth complements the study of large bones. Discovered in 2019, femur ANG19-7000 measures two metres long.

Enlarge each plate to examine labels, specimen identifiers and scale bars.

Scientific research on this topic

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