Publications and findings
Research · Publications
Dated findings trace the development of knowledge.
Explore research on Angeac-Charente and regional comparisons. Each entry connects the reference, authors and subjects studied. Media reports are grouped under Press.
30 references
Articles, theses, chapters and conference contributions are identified separately. Conference abstracts report preliminary work; several subsequently led to full papers. Online versions and repository copies of the same article are grouped together.
Scientific articles
2026 · Article · Journal of Anatomy
Heterodonty in turiasaur teeth
Heterodonty in the teeth of the turiasaurian sauropod from Angeac-Charente (Early Cretaceous, southwestern France): Comparison of anatomical and geometric morphometric approaches
Anatomical and morphometric comparison of 167 isolated teeth: five morphotypes are related to their positions in the jaw.
Clémence Le Gouellec, Ronan Allain, Jean Goedert, Romain Pintore, Jackson Carter, Rafael Royo-Torres, Dominique Augier, Raphaël Cornette
Original publication ↗ · Open-access full text ↗
Morphometrics → · Comparative anatomy and dinosaur evolution → · Sauropods →
2026 · Article · Geological Journal
How do droppings become fossils?
How Do Vertebrate Droppings Fossilise? Biogenic Versus Diagenetic Signals in Coprolites Evidenced by Elemental Mappings
Elemental mapping of 50 coprolites distinguishes biological signals from changes caused by fossilisation. Chemical composition does not simply reflect the last meal.
Kévin Rey, Ninon Robin, Pim Kaskes, Lee Rozada, Ronan Allain, Jean-Marc Baele, Elisavet Stamataki, Christophe Snoeck, Philippe Claeys
2024 · Article · Cretaceous Research, 162, 105939
Coprolites and feeding interactions
A rich coprolite assemblage from Angeac-Charente (France): A glimpse into trophic interactions within an Early Cretaceous freshwater swamp
Study of the coprolite assemblage: morphology, contents and preservation reveal feeding interactions in the wetland.
Lee Rozada, Ronan Allain, Martin Qvarnström, Kévin Rey, Romain Vullo, Jean Goedert, Dominique Augier, Ninon Robin
Coprolites → · Palaeoecology → · Crocodyliforms → · Taphonomy →
2023 · Article · eLife, 12, e83413
Femoral variation and the sexual dimorphism hypothesis
Femora from an exceptionally large population of coeval ornithomimosaurs yield evidence of sexual dimorphism in extinct theropod dinosaurs
Differences in femoral shape within a contemporaneous population are investigated as evidence of sexual dimorphism. This interpretation does not directly identify the sex of each fossil.
Romain Pintore, Raphaël Cornette, Alexandra Houssaye, Ronan Allain
Morphometrics → · Ornithomimosaurs → · Herd behaviour → · Comparative anatomy and dinosaur evolution →
2022 · Article · Geodiversitas, 44(25), 683–752
Vertebrate diversity at the Jurassic–Cretaceous transition
Vertebrate paleobiodiversity of the Early Cretaceous (Berriasian) Angeac-Charente Lagerstätte (southwestern France): implications for continental faunal turnover at the J/K boundary
Inventory of Berriasian vertebrate microremains and macroremains, with comparisons of regional continental faunas. Published counts reflect the 2022 assessment.
Ronan Allain, Romain Vullo, Lee Rozada, Jérémy Anquetin, Renaud Bourgeais, Jean Goedert, Maxime Lasseron, Jeremy Martin, Adán Pérez-García, Claire Peyre de Fabrègues, Rafael Royo-Torres, Dominique Augier, Gilles Bailly, Lilian Cazes, Yohan Despres, Auréliane Gailliègue, Bernard Gomez, Florent Goussard, Thierry Lenglet, Renaud Vacant, Mazan, Jean-François Tournepiche
Original publication ↗ · Open-access repository ↗
Diversity of life → · Comparative anatomy and dinosaur evolution → · Fish → · Turtles → · Crocodyliforms → · Pterosaurs → · Mammals → · Amphibians → · Lizards and rhynchocephalians → · Microvertebrates and screen-washing → · Stratigraphy and dating →
2021 · Article · Lethaia, 54(2), 141–165
Formation of the site: a taphonomic synthesis
A Lower Cretaceous Lagerstätte from France: a taphonomic overview of the Angeac-Charente vertebrate assemblage
Sediments, geochemistry, footprints, bone surface marks and the distribution of remains are compared to reconstruct the fossil assemblage’s history.
Lee Rozada, Ronan Allain, Romain Vullo, Jean Goedert, Dominique Augier, Amandine Jean, Jonathan Marchal, Claire Peyre de Fabrègues, Martin Qvarnström, Rafael Royo-Torres
Taphonomy → · Palaeoecology → · Ichnology → · Geochemistry → · Geology and sedimentology → · The site and its environments → · Herd behaviour →
2019 · Article · Lethaia, 52(2), 232–243 · published online in 2018, 2019 volume
A turtle shell and crocodile traces
Taphonomic study of a pleurosternid turtle shell from the Early Cretaceous of Angeac-Charente, southwest France
The study of a Pleurosternon shell analyses marks attributed to a crocodyliform, probably Goniopholis.
2018 · Article · Quaternaire, 29(1), 39–44
Trampling experiments in fine, soft sediments
Trampling experiments on bones in fine and soft sediments
Experiments on modern bones in Angeac’s sediments help explain movements, fractures and marks caused by trampling.
2018 · Article · Cretaceous Research, 90, 204–221
Pollen and spores: reassessing age and environment
Re-examination of the palynological content of the Lower Cretaceous deposits of Angeac, Charente, south-west France: Age, palaeoenvironment and taxonomic determinations
Re-examination of the palynoflora proposes a Berriasian–Valanginian age, revising the Hauterivian–Barremian estimate published in 2012.
2017 · Article · Cretaceous Research, 79, 199–213
Charophytes of the northern Aquitaine Basin
A review of the Late Jurassic–Early Cretaceous charophytes from the northern Aquitaine Basin in south-west France
Comparing Chassiron, Cherves and Angeac: charophyte assemblages provide evidence of Berriasian age and aquatic conditions.
Roch-Alexandre Benoit, Didier Néraudeau, Carles Martín-Closas
Charophytes → · Stratigraphy and dating → · Palaeoecology → · Other Charente fossil sites →
2014 · Article · Geologica Acta, 12(2), 127–135
European ornithomimosaurs: a reassessed record
European ornithomimosaurs (Dinosauria, Theropoda): an undetected record
Angeac material allows historical bones from the English Weald and the European distribution of ornithomimosaurs to be reassessed.
Ronan Allain, Romain Vullo, Jean Le Loeuff, Jean-François Tournepiche
Ornithomimosaurs → · Comparative anatomy and dinosaur evolution →
2012 · Article · Cretaceous Research, 37, 1–14
First characterisation of the Angeac fossil site
The Hauterivian–Barremian lignitic bone bed of Angeac (Charente, south-west France): stratigraphical, palaeobiological and palaeogeographical implications
The first sedimentological and palaeontological synthesis. Its Hauterivian–Barremian age was subsequently revised; the 2022 synthesis assigns the site to the Berriasian.
Didier Néraudeau, Ronan Allain, Michel Ballèvre, David J. Batten, Eric Buffetaut, Jean-Paul Colin, Marie-Pierre Dabard, Véronique Daviero-Gomez, Abderrazak El Albani, Bernard Gomez, Danièle Grosheny, Jean Le Loeuff, A. Leprince, Carles Martín-Closas, Edwige Masure, Jean-Michel Mazin, Marc Philippe, Joane Pouech, Haiyan Tong, Jean-François Tournepiche, Romain Vullo
Geology and sedimentology → · Stratigraphy and dating → · Palaeobotany → · The site and its environments → · History of the discoveries →
Thesis and chapters
2022 · Chapter · Paléontologie d’aujourd’hui, pp. 58–59
Le marécage fossile d’Angeac-Charente
A concise presentation of the fossil wetland in a multi-author palaeontology book.
2019 · Thesis · PhD · 2019MNHN0006
Taphonomy of Angeac’s vertebrate macroremains
Thesis defended at the Muséum on 4 June 2019, supervised by Nour-Eddine Jalil and Ronan Allain. It combines field data, experiments and analyses of the site.
Taphonomy → · Palaeoecology → · Ichnology → · Geochemistry →
2017 · Chapter · Dinosaures, les géants du vignoble, pp. 54–57
La taphonomie ? Élémentaire, mon cher Watson !
An educational chapter on methods and interpretations of fossil assemblages.
2017 · Chapter · Dinosaures, les géants du vignoble, pp. 128–129
Taphonomie expérimentale
An educational chapter on methods and interpretations of fossil assemblages.
2017 · Chapter · Dinosaures, les géants du vignoble, pp. 206–207
Le comportement grégaire chez les dinosaures
An educational chapter on methods and interpretations of fossil assemblages.
2017 · Chapter · Dinosaures, les géants du vignoble, pp. 220–223
Carnage à Angeac : comment sont morts les ornithomimosaures ?
An educational chapter on methods and interpretations of fossil assemblages.
Conference contributions
2026 · Conference contribution · Association paléontologique française congress, June 2026
Intraspecific and interspecific variation in Angeac-Charente sauropod teeth
A poster comparing sauropod tooth variation, presented before the August 2026 article. It is a separate conference contribution.
Clémence Le Gouellec, Jean Goedert, Alberto Cobos, Rafael Royo-Torres, Ronan Allain
Morphometrics → · Sauropods → · Comparative anatomy and dinosaur evolution →
2018 · Conference contribution · 5th Young Natural History Scientists Meeting, March 2018
Turtle shell: initial taphonomic findings
Preliminary presentation of traces on a turtle shell. The work was later developed in the Lethaia article published in the 2019 volume.
2018 · Conference contribution · APF congress, Brussels, April 2018
Taphonomic study of a pleurosternid turtle shell
APF congress contribution on the same shell study, retained as a separate stage of dissemination from the journal article.
2013 · Conference contribution · 6th International Symposium on Dinosaurs and Their Environment, Salas de los Infantes, 2013
Comparing European and Asian dinosaurs: Angeac and Laos
A comparative contribution on the Savannakhet and Angeac faunas. The reference appears in the bibliography of the 2022 synthesis, available through the link below.
Ronan Allain, Lee Rozada, Romain Vullo
Comparative anatomy and dinosaur evolution → · Ornithomimosaurs →
2023 · Conference contribution · Goldschmidt 2023
Chemical characterisation of Angeac coprolites
Conference abstract presenting chemical analyses of coprolites. Read alongside the full article published in 2026.
Kévin Rey, Ninon Robin, Pim Kaskes, Lee Rozada, Ronan Allain, Jean-Marc Baele, Elisavet Stamataki, Philippe Claeys
2018 · Conference contribution · 5th International Palaeontological Congress, Paris, July 2018
An integrated study of the Angeac bonebed
A contribution on reconstructing ecology and behaviour from the fossil assemblage.
2017 · Conference contribution · 13th Ecology and Behaviour Meeting, June 2017
Inferring past ecology and behaviour
At Angeac, bone marks, coprolites and fossil associations provide evidence of relationships between organisms.
2017 · Conference contribution · APF congress, Dijon, March 2017
Palynological re-examination of Angeac sediments
Poster announcing the palynological re-examination developed in the 2018 Cretaceous Research article.
2016 · Conference contribution · 4th ICAZ Taphonomy Working Group, Paris, September 2016
A trampling experiment in a wetland setting
An experimental presentation preceding the 2018 Quaternaire article.
2014 · Conference contribution · Society of Vertebrate Paleontology, 2014, pp. 217–218
Taphonomy of the ornithomimosaur assemblage
Presentation of initial taphonomic observations and the 2013 trampling experiment. Preliminary findings reflecting the knowledge available at that date.
Lee Rozada, Ronan Allain, Romain Vullo, Amélie Leprince, Jean-François Tournepiche
2014 · Conference contribution · 1st Young Natural History Scientists Meeting, Paris, February 2014
An ornithomimosaur herd at Angeac
Preliminary contribution on the structure and preservation of the ornithomimosaur assemblage.
2011 · Conference contribution · Society of Vertebrate Paleontology, 2011, p. 61
An assemblage dominated by ornithomimosaurs
The first scientific contribution on the abundance of ornithomimosaur bones and their taphonomic context.
Ronan Allain, Romain Vullo, Amélie Leprince, Didier Néraudeau, Jean-François Tournepiche
Ornithomimosaurs → · Herd behaviour → · History of the discoveries →
